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Table 1.

Metabolic and behavioral variables

SymbolDescriptionUnits
O2,max Maximum O2 in forced exercise (1 min) ml O2 min–1 
ADMR Average daily metabolic rate kJ day–1 
RMR Minimal resting metabolic rate (10 min) ml O2 min–1 
O2Maximum voluntary O2 averaged over 1 min ml O2 min–1 
O2Maximum voluntary O2 averaged over 2 min ml O2 min–1 
O2Maximum voluntary O2 averaged over 5 min ml O2 min–1 
Slope Slope of speed versus V̇O2 regression ml O2 kg–1 km–1 
Intercept Intercept of speed versus V̇O2 regression ml O2 kg–1 min–1 
trun Time spent running per day min day–1 
drun Distance run per day km day–1 
Vmean Mean voluntary running speed km h–1 
Vmax Maximum voluntary running speed (1.5s) km h–1 
VmaxMaximum voluntary running speed (1 min) km h–1 
VmaxMaximum voluntary running speed (2 min) km h–1 
VmaxMaximum voluntary running speed (5 min) km h–1 
Nbout Number of running bouts per day  
Boutmean Mean running bout duration 
Boutmax Maximum running bout duration 
SymbolDescriptionUnits
O2,max Maximum O2 in forced exercise (1 min) ml O2 min–1 
ADMR Average daily metabolic rate kJ day–1 
RMR Minimal resting metabolic rate (10 min) ml O2 min–1 
O2Maximum voluntary O2 averaged over 1 min ml O2 min–1 
O2Maximum voluntary O2 averaged over 2 min ml O2 min–1 
O2Maximum voluntary O2 averaged over 5 min ml O2 min–1 
Slope Slope of speed versus V̇O2 regression ml O2 kg–1 km–1 
Intercept Intercept of speed versus V̇O2 regression ml O2 kg–1 min–1 
trun Time spent running per day min day–1 
drun Distance run per day km day–1 
Vmean Mean voluntary running speed km h–1 
Vmax Maximum voluntary running speed (1.5s) km h–1 
VmaxMaximum voluntary running speed (1 min) km h–1 
VmaxMaximum voluntary running speed (2 min) km h–1 
VmaxMaximum voluntary running speed (5 min) km h–1 
Nbout Number of running bouts per day  
Boutmean Mean running bout duration 
Boutmax Maximum running bout duration 

ADMR was computed from mean daily O2 assuming 20.1 J ml–1 O2

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