... to typical anguilliform locomotion (but without head yaw) push the head into the sand; second, more pronounced undulations of the aquatic portion of the body push most of the animal below ground; third, the remaining above-ground portion of the body ceases undulation and the subterranean portion takes over...

... dihedral that may reduce rolling moments, and above 2.0 L s -1 they remain completely adducted. e-mail: [email protected] 14 6 2002 © The Company of Biologists Limited 2002 2002 needlefish Strongylura marina anguilliform locomotion steady swimming pectoral fin kinematics...