It has been shown that many factors are involved in determining the metabolic rate of an individual fish (Fry, 1957; Hughes & Shelton, 1962). The rate of respiration, for example, is influenced not only by environmental factors such as temperature and the concentration of dissolved oxygen and carbon dioxide, but also by such complicated physiological and ecological features as the size, activity and history of the fish, and external stimuli, e.g. light and sound.

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