Free-living animals differ widely in their tolerance to the absence of molecular oxygen (anoxia): most die within hours or days under these conditions, and even well-adapted forms such as intertidal invertebrates do not survive more than a month or so of continuous anoxia (Hochachka, 1980; Hochachka and Guppy, 1987; Bryant, 1991). A striking exception is the brine shrimp, Artemia franciscana, whose encysted embryos (cysts) have previously been reported to survive 4 months of continuous anoxia with no decrease in viability (Dutrieu and Chrestia-Blanchine, 1966) and 7 months with only a 25% reduction (Stocco et al. 1972). Both studies were terminated at the times given so that anoxic survival limits of this remarkable organism remain to be determined. In view of these observations, and the interest in cyst metabolism during anoxia (see Hand and Gnaiger, 1988; Clegg and Jackson, 1989a...

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