Robyn Branicky and Seigfried Hekimi Development132, 4999-5009.

There is an error in the ePress version of this article published on 19 October 2005.

On p. 5002, the sub-heading`dsc-1 encodes a Q50 Paired-like (or Prd-like) homeodomain protein'should have been the first line of the paragraph that follows it, not a sub-heading.

This error has been corrected in both the published online and print versions.

We apologise to the authors and readers for this mistake.