Dorsoventral patterning in the Drosophila embryo is controlled by the transcription factor Dorsal. A Dorsal gradient in the early embryo initiates the differentiation of several embryonic tissues by regulating target gene expression in a concentration-dependent manner. On p. 2387, Markstein and colleagues have computationally identified enhancer sequences in Dorsal target genes that are activated by intermediate levels of Dorsal, and reveal a regulatory code for neurogenic gene expression in Drosophila. Co-regulated enhancers in the rhomboid, ventral nervous system defective and brinker genes contain binding sites for Dorsal,and for the transcription factors Twist and Suppressor of Hairless, plus an additional shared motif. A search of the Drosophila genome for other enhancers with these motifs revealed two more (in vein and single-minded) that direct gene expression to the ventral neurogenic ectoderm. The same regulatory code also acts as a neurogenic enhancer in Anopheles.
Enhancing developmental patterns
Enhancing developmental patterns. Development 15 May 2004; 131 (10): e1003. doi:
Download citation file:
Sign in
Client Account
Sign in via your institution
Sign in via ShibbolethAdvertisement
Cited by
History of our journals

As our publisher, The Company of Biologists, turns 100 years old, read about Development’s journey and highlights from some its first issues, and explore the history of each of our sister journals: Journal of Cell Science, Journal of Experimental Biology, Disease Models & Mechanisms and Biology Open.
Call for papers – Lifelong Development: the Maintenance, Regeneration and Plasticity of Tissues

Development invites you to submit your latest research to our upcoming special issue – Lifelong Development: the Maintenance, Regeneration and Plasticity of Tissues. This issue will be coordinated by Guest Editors Meritxell Huch (Max Planck Institute of Molecular Cell Biology and Genetics, Germany) and Mansi Srivastava (Harvard University and Museum of Comparative Zoology, USA), working alongside our team of academic Editors. Submit your articles by 15 May 2025.
A case for broadening our view of mechanism in developmental biology

In this Perspective, B. Duygu Özpolat and colleagues survey researchers on their views on what it takes to infer mechanism in developmental biology. They examine what factors shape our idea of what we mean by ‘mechanism’ and suggest a path forward that embraces a broad outlook on the diversity of studies that advance knowledge in our field.
In preprints
Did you know that Development publishes perspectives on recent preprints? These articles help our readers navigate the ever-growing preprint literature. We welcome proposals for ‘In preprints’ articles, so please do get in touch if you’d like to contribute.