The field of evolutionary developmental (evo-devo) biology is populated with scientists from diverse disciplines who are seeking the answers to questions that have been asked for millennia: why do organisms look different,how are these organisms built, are there common themes to how they are built,are there rules governing the evolution of organismal form? The surprise of evo-devo over the last few decades is that a reductionist approach, molecular developmental genetics, has repeatedly determined that the same genes are involved in building divergent organisms. One might decide therefore that morphological evolution is now understood: it is `simply' the evolution of regulation of a conserved `tool box' of genes combined with rare but important events like gene duplication and changes in protein function. It is easy to get caught up in recent progress towards explaining diversity at the molecular level, and to forget, therefore, how much of diversity remains unexplained and how...

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