This Correction relates to Biol. Open 2013, 10:981–989.

Unfortunately, there were errors in the first published version of this article. These errors are detailed below and the original article has been changed accordingly.

The statements regarding funding were incomplete and the following text has now been included in the Acknowledgements section:

D.R. is supported by the Academy of Finland, The Sigrid Jusélius Foundation and Helsinki University Central Hospital Research Foundation.

We apologise for any confusion caused.

This is an Open Access article distributed under the terms of the Creative Commons Attribution License (, which permits unrestricted use, distribution and reproduction in any medium provided that the original work is properly attributed.